Original reference as:
- Sato, A. & N. Takezaki, H. Tichy, F. Figueroa, W.E. Mayer, J. Klein. 2003. "Origin and speciation of haplochromine fishes in East African crater lakes investigated by the analysis of their mtDNA, Mhc genes, and SINEs". Molecular Biology and Evolution. 20(9):1448–1462 (crc05436) (abstract)
Conservation: Haplochromis sp. 'lutotu' is not evaluated by the international union for the conservation of nature in the iucn red list of threatened species.